The sun, the Moon and the Sandhopper

The orientation of sandhoppers, along with that of starlings and bees, constitutes one of the classical examples of the use of chronometrically compensated celestial orienting cues. It was also the first known case of unidirectional astronomical orientation in the littoral environment (see Pardi and Papi 1952, 1953; Pardi 1957, Pardi and Ercolini 1986).